Horse Conch: Family Fasciolariidae

Horse Conch or Florida Horse Conch: Family Fasciolariidae
Triplofusus papillosus (Sowerby I, 1825)

Florida’s state shell is the Horse Conch, Triplofusus giganteus. The shell is rare on S. Hutchinson Island, I've only found one whole shell (see below).


Florida Horse Conch (S. Hutchinson Island, November 2020)

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Triplofusus papillosus
From Wikipedia, the free encyclopedia

Scientific classification
Kingdom:     Animalia
Phylum:     Mollusca
Class:     Gastropoda
Subclass:     Caenogastropoda
Order:     Neogastropoda
Family:     Fasciolariidae
Genus:     Triplofusus
Species:     T. papillosus
Binomial name
Triplofusus papillosus
(Sowerby I, 1825)

Synonyms
    Triplofusus giganteus (Kiener, 1840)
    Fasciolaria papillosa auct.
    Fasciolaria papillosa f. elongata Strebel, 1911
    Fasciolaria papillosa f. reevei Jonas, 1850
    Fasciolaria papillosa juvenis Strebel, 1911
    Fasciolaria reevei Jonas, 1850
    Pleuroploca gigantea Kiener, 1840
    Triplofusus papillosusSowerby I

Triplofusus papillosus, previously known as Pleuroploca gigantea, common name the Florida horse conch, is a species of extremely large predatory subtropical and tropical sea snail, a marine gastropod mollusc in the family Fasciolariidae, the spindle snails, tulip snails and their allies.

Although known as a horse conch, this is not a true conch, as it is not in the family Strombidae.

With a shell length that can reach 60 cm, this species is the largest gastropod in United States waters,[3] and one of the largest gastropods in the world.

Distribution
This large sea snail is found along the Atlantic coast of the Americas from the U.S. state of North Carolina to the north, to Yucatán in the Gulf of Mexico to the south.[3]
Anatomy

The animal can retract the soft parts entirely into the shell and close it with the operculum. The soft parts are bright orange in color.
Shell description
Triplofusus papillosus at the National Museum (Prague)

This species shell length can reach 24 inches (60 cm).[3]

The outline of the shell is somewhat fusiform, with a long siphonal canal, and having up to 10 whorls.[3] Its sculpture present several spiral cords and axial ribs, some of which can form knobs on the whorls' shoulders.[3]

The shell color is bright orange in very young individuals. The shell often becomes greyish white to salmon-orange when adult, with a light tan or dark brown periostracum.[3]
Ecology
A horse conch feeding on the queen conch, Lobatus gigas, in Dry Tortugas National Park, Florida
Habitat

This species dwells on sand, weed and mud flats from the low intertidal to shallow subtidal zones,[3] in 20 foot (6 m) deep water.
Feeding habits

Triplofusus papillosus is a carnivorous predatory species, and feeds on other large marine gastropods, including the tulip shell (Fasciolaria tulipa), the lightning whelk (Sinistrofulgur perversum), and the queen conch (Lobatus gigas) as well as some Murex species.[4][5] It may also present cannibalistic behaviour, feeding on smaller conspecific individuals.[4] It has been observed (in an aquarium setting) to eat small hermit crabs of the species Clibanarius vittatus.
Parasites

Parasites of Triplofusus papillosus include trematode Lophotaspis vallei.[6]
Human use
Modern times

The U.S. state of Florida declared it the state seashell in 1969. The shell is popular with shell collectors partly because of its great size.
Archaeological and anthropological uses

In classic Mayan art, the Horse Conch is shown being utilised in many ways including as paint and ink holders for elite scribes, and also as a bugle or trumpet.

In southern Florida, Native Americans, including the Calusa and Tequesta, used the horse conch to make several types of artifact. The whole shell, or more commonly only the columella, was attached to a wooden handle and used as a hammer or woodworking tool. The body whorl was used as a drinking cup. The columella was also used to make plummets or sinkers.
References

Marshall, B. (2017). Triplofusus papillosus (G. B. Sowerby I, 1825). In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=994316 on 2017-11-20
Rosenberg, G., F. Moretzsohn, and E. F. García. 2009. Gastropoda (Mollusca) of the Gulf of Mexico, Pp. 579–699 in Felder, D.L. and D.K. Camp (eds.), Gulf of Mexico–Origins, Waters, and Biota. Biodiversity. Texas A&M Press, College Station, Texas.
Leal, J.H. (2002). Gastropods. p. 99-147. In: Carpenter, K.E. (ed.). The living marine resources of the Western Central Atlantic. Volume 1: Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes, and chimaeras. FAO Species Identification Guide for Fishery Purposes and American Society of Ichthyologists and Herpetologists Special Publication No. 5. 1600p. PDF
Wells F. E., Walker D. I. & Jones D. S. (eds.) (2003). Food of giants – field observations on the diet of Syrinx aruanus (Linnaeus, 1758) (Turbinellidae) the largest living gastropod. The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth.
Toller, W.; Lewis, K-A. (2003). Queen Conch Strombus gigas (PDF). U.S.V.I. Animal Fact Sheet. 19. U.S.V.I. Department of Planning and Natural Resources Division of Fish and Wildlife. Archived from the original (PDF) on 2011-08-16.

    Alevs, Philippe V.; Vieira, Fabiano M.; Santos, Cláudia P.; Scholz, Tomáš; Luque, José L. (2015-02-12). "A Checklist of the Aspidogastrea (Platyhelminthes: Trematoda) of the World". Zootaxa. 3918 (3): 339–96. doi:10.11646/zootaxa.3918.3.2. ISSN 1175-5334. PMID 25781098.
    Rosenberg, G. 1992. Encyclopedia of Seashells. Dorset: New York. 224 pp. page(s): 91
    Rosenberg, G., F. Moretzsohn, and E. F. García. 2009. Gastropoda (Mollusca) of the Gulf of Mexico, Pp. 579–699 in Felder, D.L. and D.K. Camp (eds.), Gulf of Mexico–Origins, Waters, and Biota. Biodiversity. Texas A&M Press, College Station, Texas.
    Snyder M.A., Vermeij G.J. & Lyons W.G. (2012) The genera and biogeography of Fasciolariinae (Gastropoda, Neogastropoda, Fasciolariidae). Basteria 76(1-3): 31-70


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